Animal Ecology (Zoology 441)
Practice problems -- population growth
  1. What are the assumptions of exponential growth?  To what situations, generally, does it best apply?
  2. Four deer are introduced to an island on which there were previously no deer.  The resources for deer are excellent and the population grows at an instantaneous per capita rate of 0.2 per year.  It has been shown that if the deer population on the island exceeds 500 then the deer population will deplete all the resources and go extinct.  If the deer population keeps growing at the same per capita rate, how many years will it take before the population reaches 500 individuals?
  3. If r = 0.3 per month how long will it take for a population to double in size?
  4. You are studying the growth of a population of fruit flies living in the garbage can in your kitchen. Last week, there were 100 flies in your population.  This week, there are 1,000 flies. (a) What is the finite per capita rate of increase R? (b) What is the instantaneous per capita rate of increase r? (c) How many flies will there be one week from now? (d) What is doubling time for this population? (e) Isn't it about time to take the trash out?
  5. A species of finch has been introduced to the big island of Hawaii and the population is growing exponentially. Genetic analysis suggests that all the finches on the island were derived from a single pair of birds.  You want to know when these birds were introduced to the island.  You observe that one year, there are 500 finches.  The next year, there are 600 finches.  Assuming the per capita instantaneous growth rate has been constant, how many years ago were finches introduced to the big island?
  6. Reindeer are introduced to a nature reserve where there are no predators and no hunting is allowed.  You observe the following change in population number over five years:
  7. year number of reindeer
    1 6
    2 8
    3 11
    4 14
    5 19

    Make the appropriate graph to most clearly test whether the population is growing exponentially.  Is it?  If so, estimate r from the graph.

  8. What is density dependence? What part of the logistic growth equation makes it density dependent? Demonstrate mathematically, by using appropriate values in the logistic growth equation, that this part of the equation makes it density dependent.  What ecological phenomenon causes density dependence?
  9. A population is growing according to the simple model of logistic growth with a carrying capacity of 648.  At what population size is the population growth rate maximum?  Explain why, either in words or with a mathematical derivation.  At what population size is per capita growth rate the highest?  Why?
  10. A population of rabbits grows according to the logistic equation with a time lag of 2 time units.  The population cycles regularly. (a) Explain how a time lag can lead to population cycling. (b) How might each of the following changes in the population affect the pattern of population growth: (i) A new kind of plant, which the rabbits eat, colonizes the area.  This plant re-grows after being eaten much faster than do the plants that were originally in the area. (ii) The rabbits evolve a slower reproductive rate which lowers the value of rm for the population.
  11. What value would you assign to theta (i.e. greater than 1, 1, or less than 1) for each of the following graphs? Explain in words what each graph means with respect to the intensity of intraspecific competition at different population densities.
  12.  A population of trout in a lake has a carrying capacity of 200 and a maximum per capita growth rate of 0.75.  If the population grows according to the simple logistic growth model, how many fish should be harvested from the lake when the trout are at carrying capacity in order to maintain the population at its maximum growth rate?  Would the number of fish that should be harvested be larger, smaller or the same under any of the following conditions: (a) if the population grows according to a modified form of the logistic with a theta value of 1.5?  (b) if the maximum per capita growth rate were 0.25?  (c) If K were increased to 300?
  13. Explain in words what is meant by each of the following: rm, K, t, q, N, dN/dt, dN/Ndt, t.
  14. You study population growth in a laboratory population of Daphnia ("water fleas"; small aquatic crustaceans).  You start a population in a jar with 2 individuals, and count individuals in the jar each week, to obtain the following data:
  15.  Week # Population Size
    2
    2 4
    6
    10
    5 16
    6 24
    7 32
    37
    39
    10  40
    11  40
    12  40

    Plot population size versus time. Which version of logistic growth (e.g. simple, with time lag, non-linear...) does this population appear to fit? Use the graph to estimate K. Finally, estimate rm (to do so, remember that initially a population undergoing logistic growth will grow exponentially).

  16. What is the traditional definition of "carrying capacity" as used for most animal species, and, according to Barrett and Odum (2000), why does it not apply well to humans?  What must be considered in addition to density to develop a meaningful definition of the maximum carrying capacity for humans?
  17. Barrett and Odum (2000) state that the "overshoot pattern" occurs in nature and in human affairs.  What is the "overshoot pattern"?  What are examples of causes for the overshoot pattern in nature?  What are causes of the overshoot pattern in human affairs?  How does the "ecological footprint" or a city such as Vancouver BC provide evidence of an overshoot pattern?
  18. According to Barrett and Odum (2000), what is the difference between economic development and economic growth?  What is the ecological argument for focusing on economic development rather than economic growth?
  19. What is the economic welfare threshold? Why do Barrett and Odum (2000) argue that it is equivalent to the optimum carrying capacity?
  20. What do you think of the Barrett and Odum (2000) recommendations for an integrated (ecologic/economic) capitalism?  Are they desirable?  Are they practical?  Why/why not?