Department of Biological Sciences
Biology 391: Organic Evolution
Course Information
Course description and goals: This course focuses
on evolution as studied in extant (currently living, not extinct) species.
The goals are to cover general principles of microevolution and macroevolution
and to focus on certain areas that are currently being actively researched.
Students are expected to learn to apply hypotheses and models of evolution
to novel situations, and to evaluate suppport for conflicting hypotheses.
Where relevant, evolutionary principles are applied to humans.
Prerequisites: Biology 110 and 120. Biology 336 (Genetics) is
strongly
recommended!
Textbooks and Other Readings: You are required to have the textbook,
Evolutionary Analysis, 2nd edition, by Freeman and Herron (2001).
You will be expected to print additional materials from the web to read
and bring to class. The main additional materials to print from the web
are chapters from the Evolution Lab Manual (see "Course Related Links"
below.) Additional readings will alse be required; these will be placed
on reserve in the library.
Grades are based on the following; click to obtain more information:
Grades are based on the percentage of the possible points that you obtain:
90-100% = A, 80-89%=B, 70-79%=C, 60-69%=D, <60%=F. If you make the lower
boundary for a grade level, you are guaranteed that grade. I reserve
the right to make the boundaries a little lower based on the final point
distribution
(but don't count on this; in past semesters they have not dropped more
than about 2 percentage points and sometimes do not drop at all.) There
is NO possibility of extra credit.
Course Policies: READ THIS SECTION CAREFULLY, IT'S IMPORTANT! Exams
must be taken during the scheduled times. No make-up or early exams will
be given. Your percentage on the final exam will substitute for one of
the three exams if you must miss an exam or if the substitution benefits
you. Outside assignments and homework must be completed by the deadline;
late assignments will NOT be accepted unless you have a serious (medical
or family emergency) excuse that you OK with me as soon as possible. Attendance
and participation in lab sections is required. You will not receive participation
or homework points if you do not attend; in addition,
your grade will
be dropped one letter grade for every three lab sessions you miss.
Any student eligible for and requesting academic accommodations due
to a disability is requested to provide a letter of accommodation from
P.A.C.E. or Student Academic Support Center within the first two weeks
of the semester.
Academic Honesty: Cheating and plagiarism are punishable by a
grade of F in the course; individuals caught in any form of academic honesty
can expect to be reported to the Vice Chancellor for Student Affairs. Note
that it is as dishonest (and unfair to other students) to allow someone
to copy your work as it is to copy someone else's work.
Electronic Devices: CELL PHONES MUST BE TURNED OFF IN CLASS.
(if there's an emergency situation where you have to be reachable, you
must tell me before class, and be prepared to get up and leave the
class before answering it.) Do not use other electronic devices (portable
CD players, etc.) in class, either. Anyone listening to such a device
or answering a cell phone during an exam is getting extra information and
is therefore cheating and will receive an automatic F in the course.
Information on Lab Sections: Most lab sections are discussion
sections during which you will apply your knowledge of evolution (in ways
in which you will also be expected to apply it on exams) by discussion
of study questions or assignment results. The study questions are
available in the Evolution Lab Manual web pages. We will use most
lab periods to get started working on the material in the lab manual; it
is your responsibility to make sure you can answer all questions relevant
to lectures and assigned readings, not just the ones we go through in lab.
You MUST print out the required chapter(s) and bring them with you each
week to your laboratory section. Since labs may cover questions from previous
weeks as well as the current week, you must bring all chapters through
whatever is current to lab each week. To prepare for lab, you SHOULD
print them out ahead of time and try to work on the questions that cover
the relevant lectures and reading before coming to lab; the more you prepare
in advance, the better the labs will help you to learn the material.
Some lab sessions will cover other materials (other readings, results from
assignments, or other information from the web); this is indicated on the
course schedule below.
Information on Written Assignments:
Homework assignments are questions that will
require a brief written (or occasionally mathematical or graphical) answer.
They are designed to give you practice applying the material we are learning
in the way that you will be required to do on essay questions on the exam.
They are like the questions from the lab manual and you can use them as
a guide to helping you see the level at which you are expected to know
all the lab manual questions. The answer to one homework question
is due almost every week. You must turn it in at the start
of your lab period; it must be handed in by you (you can't give it to someone
else to hand in for you). Homework assignments requiring a written
answer must be typed, double spaced. Homework assignments requiring
a mathematical answer may be typed or handwritten; in either case, they
must be clearly organized and show all formulas and intermediate steps.
Homework assignments requiring a graph or figure may be hand drawn; the
drawing must be clear and large. No late homework will
be accepted; to receive homework credit you must be in your scheduled
lab (or have arranged ahead of time with me to attend a different lab;
in which case you can turn them in at the start of that lab) on time.
You must bring two copies of each homework assignment to lab as
we will correct them in lab and you will be expected to make corrections
on your copy of the homework as originals may not be returned; because
corrections are made during lab you must attend the entire lab to receive
homework credit.
Other assignments (computer assignment, results paper, and
critique): the final version must be handed in on paper, typed, double-spaced.
Rough drafts: I STRONGLY ENCOURAGE you to submit rough
drafts of assignments. Rough drafts can be submitted typed, double
spaced on paper or via e-mail -- I recommend e-mail submissions, as I can
able to type more extensive, and more legible, comments on them than I
can scribble on a hard copy. If you absolutely MUST do so, you can
submit a handwritten rough draft (but not final copy) -- if you submit
a handwritten draft you MUST leave a blank line between each written line
and it must be legible (if I can't read it easily I will not comment on
it.) For me to have time to comment on rough drafts, I must receive
them at least three working days before the due date of the paper (i.e.for
weeks without holidays -- if the paper is due on a Friday, I need
the draft by Tuesday. If the paper is due on a Monday, I need it
by the previous Wednesday). The earlier I receive a rough draft,
the more time I'll have to comment on it and the more you can benefit.
IT IS YOUR RESPONSIBILITY to
KEEP A COPY of every assignment
you hand in (paper, homework, graphs, rough drafts...) until you receive
your returned, graded assignment; if I ever don't get something you think
you handed in, and you can produce your copy immediately, I am very likely
to accept it but if you CAN'T produce it I will not take it.
Course-related Web Links:
Course content:
the following gives the tentative
lecture schedule for Fall Semester, 2002. As lectures are
completed, they will be made available on the web; click on links to lecture
topics to see the lecture (if they're not highlighted as links, they're
not available yet.) Under each week's set of lectures, you will find
a link to the the HOMEWORK ASSIGNMENT
for
the week. EXAM times and DUE DATES FOR ASSIGNMENTS
are
also given. This lecture schedule is tentitive: I may change topics/dates
if I get behind or ahead. Exam and due dates are very unlikely to
change. Assigned readings (readings from which you are expected to
know material beyond what is covered in lecture) and lab manual chapters
with questions about the lectures are also given; in your lab manual, it
indicates what textbook chapters also cover material related to what is
covered in lecture. Readings may be modified during the semester; if they
are, I'll let you know and make changes here. If an assigned reading refers
to a chapter, it means a chapter in your textbook. If it refers to
a paper, the paper is on reserve in the library. For labs:
be sure to print out and bring with you all lab manual chapters
relevant to all lecture and reading material covered prior to the lab (including
all
chapters from previous weeks.)
Assigned readings (which contain material beyond what is covered in
lecture) are called assigned readings and printed in red. Other references
to textbook chapters indicate where you can find background and additional
reading on topics covered in lecture.
-
Week 1: 6-8 Jan Assigned Reading: Freeman
and Herron (2001) Chapter 1
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Week 2: 13-15 Jan
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Week 3: 20-22-24 Jan Assigned Readings: "Nonoverlapping
Magisteria" by S.J. Gould
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Week 4: 27-29 Jan
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Week 5: 3-5 Feb
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Monday, 3 Feb: LAB over computer assignment.
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BEFORE LAB PREPARE FOR THE FOLLOWING: During lab, each
group will be responsible for being able to answer an assigned set of the
practice questions for the graphs and tables. Here are the questions
for which you are responsible in lab:
-
Group 1: Practice Questions for Graphs 1 and 2
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Group 2: Practice Questions for Graphs 1 and 3
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Group 3: Practice Questions for Graphs 1 and 4
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Group 4: Practice Questions for Graph 1, Graph 5, and Table 2
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After completing lab, you should make sure you can
answer questions from ALL graphs and tables discussed in lab and from Table
3. You should also be sure you could recognize or draw graphs representing
any of the situation. Finally, for the exam, you will have to answer
an essay question (in addition to other questions) over this material.
This essay question will be ONE of the questions from Lab Manual Chapter
VI, so CLICK HERE
for Lab Manual Chapter VI which has essay questions you need to prepare
for exam 1 (one of the essay questions here WILL be on the first exam;
material from the other questions will also be on the exam; be sure to
prepare to answer any one of them!)
-
Click here for HOMEWORK
QUESTION 4 Due at start of lab on 3 Feb.
-
Wednesday, 5 February: Lecture: Measuring
Variation in Polygenic Traits (Lab
Manual Ch. IX) and Evolution
of Polygenic Traits (Lab
Manual Ch. IX) Freeman and Herron (2001) Chapter 7
-
Friday, 7 Feb: LAB We will focus on lab manual chapters not
yet discussed in lab, especially VII and IX, but since this is also an
exam review please bring ALL lab manual chapters associated with class
so far (I-X)
-
Click here for HOMEWORK
QUESTION 5
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Week 6: 10-12 Feb
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Week 7: 17-19 Feb
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Week 8: 24-26 Feb
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Lecture 1: Speciation:
General Principles; Genetic Analyses (Lab
Manual Ch. XIII)
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Lecture 2: Speciation (continued from previous lecture)
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You MUST get your character tables and phylogenetic tree for the Phylogenetic
Analysis Assignment checked by me by 5:00 p.m. on Friday 28 Feb or you
will not be eligible to complete the Phylogenetic Analysis Assignment.
If there are any problems with your characters or tree, you MUST show me
the corrected characters and tree by 5:00 p.m. on Mon. 3 Mar. to be allowed
to receive credit for the Phylogenetic Analysis Assignment.
-
Click here for HOMEWORK
QUESTION 8
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Week 9: 3-5 March
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Week 10: 17-19 March
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Week 11: 24-26 Mar
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Week 12: 31 Mar - 2 Apr
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Lecture 1: DNA and Phylogenetic Analysis Continued
(Lab
Manual Ch. XII)
Freeman and Herron (2001) Chapter 18
-
Lecture 2: Molecular
Evolution II: DNA and Evolutionary Processes (Lab
Manual Ch. XII)
-
BEFORE LAB: Read the "Critique
Warm-up Paper." Be able to define terms in bold-face; ASK ME AHEAD
OF TIME if you have questions. Groups 1 and 3: look for strong points
in the paper (methods that should result in a reliable phylogeny.)
Groups 2 and 4: look for weak points in the paper (methods that make the
phylogeny questionable or results that do not support the conclusion well.)
Be prepared to discuss these strong and weak points in lab! Note
that this material will help you start the critique assignment and will
be covered on Exam 3.
-
Click here for HOMEWORK
QUESTION 12
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Week 13: 7-9 Apr.
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Week 14: 14-16 Apr
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Week 15: 21-23 Apr
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Lecture 1: Lecture 1: Catch up from previous lectures (will be on exam)
and, time permitting, Human
Macroevolution (Lab
Manual Ch. XXII) (Will not be covered on exam 3; will be on final)
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Lecture 2: EXAM 3
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Week 16: 28 Apr
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Human Microevolution;
and Human Behavior
(Lab
Manual Ch. XXII) Freeman and Herron (2001) Chapter 16
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Assigned Reading: to prepare for an assigned essay
question on the final exam, you will need to read from one of the
following chapters: 14, 18, or 19 and prepare to answer
your choice
of one of the following questions from your Lab
Manual chapter XXIII: question 3 or question 4
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CRITIQUE DUE Mon 28 Apr BY 5:00 P.M. (Hand in to me in my office
OR give to the Biology Department Office)
Interesting Evolution-related Web Sites:
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An
Introduction to Evolutionary Biology, by Chris Colby. Introduces the
major areas of evolution; presents a strong (and rather strongly worded)
argument against "Creationism."
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Five
Major Misconceptions about Evolution, a site that clearly describes
five arguments that have incorrectly been used in attempts to demonstrate
that evolution does not occur, with explanations of what is wrong with
these arguments.
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Observed
instances of speciation. Descriptions of situations in which the formation
of new species has been observed.
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Transitional
Vertebrate Fossils. Description/discussion of fossils that demonstrate
transitional changes between species/groups of vertebrates.
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An explanation
of the concept of heritability
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The Tree of
Life , designed to show (ultimately) what is known about the phylogeny
of all life.
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The University of California
at Berkeley Museum of Paleontology , with excellent virtual exhibits,
clear information on the phylogeny of many groups of organisms, some good
explanations of basic evolutionary processes, and more.